MicroRNA-24-3p Targets Notch and Other Vascular Morphogens to Regulate Post-ischemic Microvascular Responses in Limb Muscles

<p dir="ltr">MicroRNAs (miRs) regulate complex processes, including angiogenesis, by targeting multiple mRNAs. miR-24-3p-3p directly represses eNOS, GATA2, and PAK4 in endothelial cells (ECs), thus inhibiting angiogenesis during development and in the infarcted heart. miR-24-3p is wi...

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محفوظ في:
التفاصيل البيبلوغرافية
المؤلف الرئيسي: Micol Marchetti (18623410) (author)
مؤلفون آخرون: Marco Meloni (7714451) (author), Maryam Anwar (6458852) (author), Ayman Al-Haj-Zen (18623413) (author), Graciela Sala-Newby (18623416) (author), Sadie Slater (18623419) (author), Kerrie Ford (18623422) (author), Andrea Caporali (7714448) (author), Costanza Emanueli (299596) (author)
منشور في: 2020
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_version_ 1864513513524822016
author Micol Marchetti (18623410)
author2 Marco Meloni (7714451)
Maryam Anwar (6458852)
Ayman Al-Haj-Zen (18623413)
Graciela Sala-Newby (18623416)
Sadie Slater (18623419)
Kerrie Ford (18623422)
Andrea Caporali (7714448)
Costanza Emanueli (299596)
author2_role author
author
author
author
author
author
author
author
author_facet Micol Marchetti (18623410)
Marco Meloni (7714451)
Maryam Anwar (6458852)
Ayman Al-Haj-Zen (18623413)
Graciela Sala-Newby (18623416)
Sadie Slater (18623419)
Kerrie Ford (18623422)
Andrea Caporali (7714448)
Costanza Emanueli (299596)
author_role author
dc.creator.none.fl_str_mv Micol Marchetti (18623410)
Marco Meloni (7714451)
Maryam Anwar (6458852)
Ayman Al-Haj-Zen (18623413)
Graciela Sala-Newby (18623416)
Sadie Slater (18623419)
Kerrie Ford (18623422)
Andrea Caporali (7714448)
Costanza Emanueli (299596)
dc.date.none.fl_str_mv 2020-03-03T03:00:00Z
dc.identifier.none.fl_str_mv 10.3390/ijms21051733
dc.relation.none.fl_str_mv https://figshare.com/articles/journal_contribution/MicroRNA-24-3p_Targets_Notch_and_Other_Vascular_Morphogens_to_Regulate_Post-ischemic_Microvascular_Responses_in_Limb_Muscles/25912141
dc.rights.none.fl_str_mv CC BY 4.0
info:eu-repo/semantics/openAccess
dc.subject.none.fl_str_mv Biological sciences
Biochemistry and cell biology
Biomedical and clinical sciences
Cardiovascular medicine and haematology
miR-24-3p
angiogenesis
endothelial cells
Notch
β-catenin
limb ischemia
dc.title.none.fl_str_mv MicroRNA-24-3p Targets Notch and Other Vascular Morphogens to Regulate Post-ischemic Microvascular Responses in Limb Muscles
dc.type.none.fl_str_mv Text
Journal contribution
info:eu-repo/semantics/publishedVersion
text
contribution to journal
description <p dir="ltr">MicroRNAs (miRs) regulate complex processes, including angiogenesis, by targeting multiple mRNAs. miR-24-3p-3p directly represses eNOS, GATA2, and PAK4 in endothelial cells (ECs), thus inhibiting angiogenesis during development and in the infarcted heart. miR-24-3p is widely expressed in cardiovascular cells, suggesting that it could additionally regulate angiogenesis by acting on vascular mural cells. Here, we have investigated: (1) new miR-24-3p targets; (2) the expression and the function of miR-24-3p in human vascular ECs; (3) the impact of miR-24-3p inhibition in the angiogenesis reparative response to limb ischemia in mice. Using bioinformatics target prediction platforms and 3′-UTR luciferase assays, we newly identified Notch1 and its Delta-like ligand 1 (Dll1) to be directly targeted by miR-24-3p. miR-24-3p was expressed in human ECs and pericytes cultured under normal conditions. Exposure to hypoxia increased miR-24-3p in ECs but not in pericytes. Transfection with a miR-24-3p precursor (pre-miR-24-3p) increased miR-24-3p expression in ECs, reducing the cell survival, proliferation, and angiogenic capacity. Opposite effects were caused by miR-24-3p inhibition. The anti-angiogenic action of miR-24-3p overexpression could be prevented by simultaneous adenovirus (Ad)-mediated delivery of constitutively active Notch intracellular domain (NICD) into cultured ECs. We next demonstrated that reduced Notch signalling contributes to the anti-angiogenic effect of miR-24-3p in vitro. In a mouse unilateral limb ischemia model, local miR-24-3p inhibition (by adenovirus-mediated miR-24-3p decoy delivery) restored endothelial Notch signalling and increased capillary density. However, the new vessels appeared disorganised and twisted, worsening post-ischemic blood perfusion recovery. To better understand the underpinning mechanisms, we widened the search for miR-24-3p target genes, identifying several contributors to vascular morphogenesis, such as several members of the Wingless (Wnt) signalling pathway, β-catenin signalling components, and VE-cadherin, which synergise to regulate angiogenesis, pericytes recruitment to neoformed capillaries, maturation, and stabilization of newly formed vessels. Among those, we next focussed on β-catenin to demonstrate that miR-24-3p inhibition reduces β-catenin expression in hypoxic ECs, which is accompanied by reduced adhesion of pericytes to ECs. In summary, miR-24-3p differentially targets several angiogenesis modulators and contributes to autonomous and non-autonomous EC crosstalk. In ischemic limbs, miR-24-3p inhibition increases the production of dysfunctional microvessels, impairing perfusion. Caution should be observed in therapeutic targeting of miR-24-3p.</p><h2>Other Information</h2><p dir="ltr">Published in: International Journal of Molecular Sciences<br>License: <a href="https://creativecommons.org/licenses/by/4.0/" target="_blank">https://creativecommons.org/licenses/by/4.0/</a><br>See article on publisher's website: <a href="https://dx.doi.org/10.3390/ijms21051733" target="_blank">https://dx.doi.org/10.3390/ijms21051733</a></p>
eu_rights_str_mv openAccess
id Manara2_05130abcc705bf3438d4ba7146bc971c
identifier_str_mv 10.3390/ijms21051733
network_acronym_str Manara2
network_name_str Manara2
oai_identifier_str oai:figshare.com:article/25912141
publishDate 2020
repository.mail.fl_str_mv
repository.name.fl_str_mv
repository_id_str
rights_invalid_str_mv CC BY 4.0
spelling MicroRNA-24-3p Targets Notch and Other Vascular Morphogens to Regulate Post-ischemic Microvascular Responses in Limb MusclesMicol Marchetti (18623410)Marco Meloni (7714451)Maryam Anwar (6458852)Ayman Al-Haj-Zen (18623413)Graciela Sala-Newby (18623416)Sadie Slater (18623419)Kerrie Ford (18623422)Andrea Caporali (7714448)Costanza Emanueli (299596)Biological sciencesBiochemistry and cell biologyBiomedical and clinical sciencesCardiovascular medicine and haematologymiR-24-3pangiogenesisendothelial cellsNotchβ-cateninlimb ischemia<p dir="ltr">MicroRNAs (miRs) regulate complex processes, including angiogenesis, by targeting multiple mRNAs. miR-24-3p-3p directly represses eNOS, GATA2, and PAK4 in endothelial cells (ECs), thus inhibiting angiogenesis during development and in the infarcted heart. miR-24-3p is widely expressed in cardiovascular cells, suggesting that it could additionally regulate angiogenesis by acting on vascular mural cells. Here, we have investigated: (1) new miR-24-3p targets; (2) the expression and the function of miR-24-3p in human vascular ECs; (3) the impact of miR-24-3p inhibition in the angiogenesis reparative response to limb ischemia in mice. Using bioinformatics target prediction platforms and 3′-UTR luciferase assays, we newly identified Notch1 and its Delta-like ligand 1 (Dll1) to be directly targeted by miR-24-3p. miR-24-3p was expressed in human ECs and pericytes cultured under normal conditions. Exposure to hypoxia increased miR-24-3p in ECs but not in pericytes. Transfection with a miR-24-3p precursor (pre-miR-24-3p) increased miR-24-3p expression in ECs, reducing the cell survival, proliferation, and angiogenic capacity. Opposite effects were caused by miR-24-3p inhibition. The anti-angiogenic action of miR-24-3p overexpression could be prevented by simultaneous adenovirus (Ad)-mediated delivery of constitutively active Notch intracellular domain (NICD) into cultured ECs. We next demonstrated that reduced Notch signalling contributes to the anti-angiogenic effect of miR-24-3p in vitro. In a mouse unilateral limb ischemia model, local miR-24-3p inhibition (by adenovirus-mediated miR-24-3p decoy delivery) restored endothelial Notch signalling and increased capillary density. However, the new vessels appeared disorganised and twisted, worsening post-ischemic blood perfusion recovery. To better understand the underpinning mechanisms, we widened the search for miR-24-3p target genes, identifying several contributors to vascular morphogenesis, such as several members of the Wingless (Wnt) signalling pathway, β-catenin signalling components, and VE-cadherin, which synergise to regulate angiogenesis, pericytes recruitment to neoformed capillaries, maturation, and stabilization of newly formed vessels. Among those, we next focussed on β-catenin to demonstrate that miR-24-3p inhibition reduces β-catenin expression in hypoxic ECs, which is accompanied by reduced adhesion of pericytes to ECs. In summary, miR-24-3p differentially targets several angiogenesis modulators and contributes to autonomous and non-autonomous EC crosstalk. In ischemic limbs, miR-24-3p inhibition increases the production of dysfunctional microvessels, impairing perfusion. Caution should be observed in therapeutic targeting of miR-24-3p.</p><h2>Other Information</h2><p dir="ltr">Published in: International Journal of Molecular Sciences<br>License: <a href="https://creativecommons.org/licenses/by/4.0/" target="_blank">https://creativecommons.org/licenses/by/4.0/</a><br>See article on publisher's website: <a href="https://dx.doi.org/10.3390/ijms21051733" target="_blank">https://dx.doi.org/10.3390/ijms21051733</a></p>2020-03-03T03:00:00ZTextJournal contributioninfo:eu-repo/semantics/publishedVersiontextcontribution to journal10.3390/ijms21051733https://figshare.com/articles/journal_contribution/MicroRNA-24-3p_Targets_Notch_and_Other_Vascular_Morphogens_to_Regulate_Post-ischemic_Microvascular_Responses_in_Limb_Muscles/25912141CC BY 4.0info:eu-repo/semantics/openAccessoai:figshare.com:article/259121412020-03-03T03:00:00Z
spellingShingle MicroRNA-24-3p Targets Notch and Other Vascular Morphogens to Regulate Post-ischemic Microvascular Responses in Limb Muscles
Micol Marchetti (18623410)
Biological sciences
Biochemistry and cell biology
Biomedical and clinical sciences
Cardiovascular medicine and haematology
miR-24-3p
angiogenesis
endothelial cells
Notch
β-catenin
limb ischemia
status_str publishedVersion
title MicroRNA-24-3p Targets Notch and Other Vascular Morphogens to Regulate Post-ischemic Microvascular Responses in Limb Muscles
title_full MicroRNA-24-3p Targets Notch and Other Vascular Morphogens to Regulate Post-ischemic Microvascular Responses in Limb Muscles
title_fullStr MicroRNA-24-3p Targets Notch and Other Vascular Morphogens to Regulate Post-ischemic Microvascular Responses in Limb Muscles
title_full_unstemmed MicroRNA-24-3p Targets Notch and Other Vascular Morphogens to Regulate Post-ischemic Microvascular Responses in Limb Muscles
title_short MicroRNA-24-3p Targets Notch and Other Vascular Morphogens to Regulate Post-ischemic Microvascular Responses in Limb Muscles
title_sort MicroRNA-24-3p Targets Notch and Other Vascular Morphogens to Regulate Post-ischemic Microvascular Responses in Limb Muscles
topic Biological sciences
Biochemistry and cell biology
Biomedical and clinical sciences
Cardiovascular medicine and haematology
miR-24-3p
angiogenesis
endothelial cells
Notch
β-catenin
limb ischemia