A human infertility-associated KASH5 variant promotes mitochondrial localization

<p>KASH5 is the most recently identified member of the KASH domain family of tail anchored, outer nuclear membrane (ONM) and endoplasmic reticulum (ER) proteins. During meiosis prophase I, KASH5 and SUN1 form a complex that spans the nuclear envelope and which links the telomeres of meiotic ch...

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Main Author: Sana A. Bentebbal (14152929) (author)
Other Authors: Bakhita R. Meqbel (14152932) (author), Anna Salter (14152935) (author), Victoria Allan (7890212) (author), Brian Burke (248053) (author), Henning F. Horn (14152938) (author)
Published: 2022
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author Sana A. Bentebbal (14152929)
author2 Bakhita R. Meqbel (14152932)
Anna Salter (14152935)
Victoria Allan (7890212)
Brian Burke (248053)
Henning F. Horn (14152938)
author2_role author
author
author
author
author
author_facet Sana A. Bentebbal (14152929)
Bakhita R. Meqbel (14152932)
Anna Salter (14152935)
Victoria Allan (7890212)
Brian Burke (248053)
Henning F. Horn (14152938)
author_role author
dc.creator.none.fl_str_mv Sana A. Bentebbal (14152929)
Bakhita R. Meqbel (14152932)
Anna Salter (14152935)
Victoria Allan (7890212)
Brian Burke (248053)
Henning F. Horn (14152938)
dc.date.none.fl_str_mv 2022-11-22T21:17:18Z
dc.identifier.none.fl_str_mv 10.1038/s41598-021-89439-2
dc.relation.none.fl_str_mv https://figshare.com/articles/journal_contribution/A_human_infertility-associated_KASH5_variant_promotes_mitochondrial_localization/21598233
dc.rights.none.fl_str_mv CC BY 4.0
info:eu-repo/semantics/openAccess
dc.subject.none.fl_str_mv Reproductive medicine
Multidisciplinary
dc.title.none.fl_str_mv A human infertility-associated KASH5 variant promotes mitochondrial localization
dc.type.none.fl_str_mv Text
Journal contribution
info:eu-repo/semantics/publishedVersion
text
contribution to journal
description <p>KASH5 is the most recently identified member of the KASH domain family of tail anchored, outer nuclear membrane (ONM) and endoplasmic reticulum (ER) proteins. During meiosis prophase I, KASH5 and SUN1 form a complex that spans the nuclear envelope and which links the telomeres of meiotic chromosomes to cytoplasmic dynein. This connection is essential for homologous chromosome dynamics and pairing. A recent study identified a variant in human KASH5 (L535Q) that correlated with male infertility associated with azoospermia. However, no molecular mechanism was described. Here, we report that this amino acid substitution, within the KASH5 transmembrane domain (TMD) has no predicted effects on secondary structure. However, the overall hydrophobicity of the L535Q TMD, is calculated to be lower than the wild-type KASH5, based on the GES (Goldman–Engelman–Steitz) amino acid hydrophobicity scale. This change in hydrophobicity profoundly affects the subcellular localization of KASH5. Through a series of amino acid substitution studies, we show that the L535Q substitution perturbs KASH5 localization to the ER and ONM and instead results in mistargeting to the mitochondria membrane. We suggest that this mislocalization accounts for the infertility and azoospermia phenotype in patients.</p><h2>Other Information</h2> <p> Published in: Scientific Reports<br> License: <a href="https://creativecommons.org/licenses/by/4.0" target="_blank">https://creativecommons.org/licenses/by/4.0</a><br>See article on publisher's website: <a href="http://dx.doi.org/10.1038/s41598-021-89439-2" target="_blank">http://dx.doi.org/10.1038/s41598-021-89439-2</a></p>
eu_rights_str_mv openAccess
id Manara2_562e096e49b76ae94086cf0d55035cc1
identifier_str_mv 10.1038/s41598-021-89439-2
network_acronym_str Manara2
network_name_str Manara2
oai_identifier_str oai:figshare.com:article/21598233
publishDate 2022
repository.mail.fl_str_mv
repository.name.fl_str_mv
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rights_invalid_str_mv CC BY 4.0
spelling A human infertility-associated KASH5 variant promotes mitochondrial localizationSana A. Bentebbal (14152929)Bakhita R. Meqbel (14152932)Anna Salter (14152935)Victoria Allan (7890212)Brian Burke (248053)Henning F. Horn (14152938)Reproductive medicineMultidisciplinary<p>KASH5 is the most recently identified member of the KASH domain family of tail anchored, outer nuclear membrane (ONM) and endoplasmic reticulum (ER) proteins. During meiosis prophase I, KASH5 and SUN1 form a complex that spans the nuclear envelope and which links the telomeres of meiotic chromosomes to cytoplasmic dynein. This connection is essential for homologous chromosome dynamics and pairing. A recent study identified a variant in human KASH5 (L535Q) that correlated with male infertility associated with azoospermia. However, no molecular mechanism was described. Here, we report that this amino acid substitution, within the KASH5 transmembrane domain (TMD) has no predicted effects on secondary structure. However, the overall hydrophobicity of the L535Q TMD, is calculated to be lower than the wild-type KASH5, based on the GES (Goldman–Engelman–Steitz) amino acid hydrophobicity scale. This change in hydrophobicity profoundly affects the subcellular localization of KASH5. Through a series of amino acid substitution studies, we show that the L535Q substitution perturbs KASH5 localization to the ER and ONM and instead results in mistargeting to the mitochondria membrane. We suggest that this mislocalization accounts for the infertility and azoospermia phenotype in patients.</p><h2>Other Information</h2> <p> Published in: Scientific Reports<br> License: <a href="https://creativecommons.org/licenses/by/4.0" target="_blank">https://creativecommons.org/licenses/by/4.0</a><br>See article on publisher's website: <a href="http://dx.doi.org/10.1038/s41598-021-89439-2" target="_blank">http://dx.doi.org/10.1038/s41598-021-89439-2</a></p>2022-11-22T21:17:18ZTextJournal contributioninfo:eu-repo/semantics/publishedVersiontextcontribution to journal10.1038/s41598-021-89439-2https://figshare.com/articles/journal_contribution/A_human_infertility-associated_KASH5_variant_promotes_mitochondrial_localization/21598233CC BY 4.0info:eu-repo/semantics/openAccessoai:figshare.com:article/215982332022-11-22T21:17:18Z
spellingShingle A human infertility-associated KASH5 variant promotes mitochondrial localization
Sana A. Bentebbal (14152929)
Reproductive medicine
Multidisciplinary
status_str publishedVersion
title A human infertility-associated KASH5 variant promotes mitochondrial localization
title_full A human infertility-associated KASH5 variant promotes mitochondrial localization
title_fullStr A human infertility-associated KASH5 variant promotes mitochondrial localization
title_full_unstemmed A human infertility-associated KASH5 variant promotes mitochondrial localization
title_short A human infertility-associated KASH5 variant promotes mitochondrial localization
title_sort A human infertility-associated KASH5 variant promotes mitochondrial localization
topic Reproductive medicine
Multidisciplinary