AKT regulates UGCG expression via epigenomic alterations.

<p>(<b>A</b>) A schematic diagram showing AKT-mediated phosphorylation of DNMT1 leading to hypomethylation of CpG islands that further enhances UGCG expression. (<b>B</b>) Immunoblot showing alteration in phosphorylation of DNMT1 by pan-phospho-ser antibody in MCF-7_RIC...

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Main Author: Mohammad Nafees Ansari (22232505) (author)
Other Authors: Somesh K. Jha (20722073) (author), Ali Khan (568900) (author), Kajal Rajput (11403230) (author), Nishant Pandey (5578880) (author), Dolly Jain (11403227) (author), Rajeshwari Tripathi (22232508) (author), Nihal Medatwal (4251403) (author), Pankaj Sharma (58141) (author), Sudeshna Datta (3643081) (author), Animesh Kar (4583296) (author), Trishna Pani (22232511) (author), Sk Asif Ali (22232514) (author), Kaushavi Cholke (19669386) (author), Kajal Rana (11403212) (author), Valiya P. Snijesh (22232517) (author), Geetashree Mukherjee (264708) (author), Suryanarayana V. S. Deo (22232520) (author), Soumen Basak (1681390) (author), Ashutosh Mishra (231926) (author), Jyothi S. Prabhu (3173547) (author), Arnab Mukhopadhyay (5645819) (author), Avinash Bajaj (1910212) (author), Ujjaini Dasgupta (7483922) (author)
Published: 2025
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Summary:<p>(<b>A</b>) A schematic diagram showing AKT-mediated phosphorylation of DNMT1 leading to hypomethylation of CpG islands that further enhances UGCG expression. (<b>B</b>) Immunoblot showing alteration in phosphorylation of DNMT1 by pan-phospho-ser antibody in MCF-7_RICTOR<sup>SH</sup> and MCF-7_ZFX<sup>OE</sup> cells compared to MCF-7 cells. (<b>C</b>) Immunoblots showing a dose-dependent decrease in pAKT and UGCG expression in MCF-7_ZFX<sup>OE</sup> cells on treatment with AKT inhibitor MK2206. (<b>D</b>) Results from qRT-PCR (mean ± SEM, <i>n</i> = 4) show a decrease in <i>UGCG</i> expression in MCF-7_ZFX<sup>OE</sup> cells on AKT inhibition by MK2206. (<b>E</b>) Immunoblot reveals a decrease in phosphorylation of DNMT1 on treatment of MCF-7_ZFX<sup>OE</sup> cells with AKT inhibitor MK2206. (<b>F</b>, <b>G</b>) Results from qRT-PCR (mean ± SEM, <i>n</i> = 3) (F) and immunoblot (G) show increased <i>UGCG</i> expression in MCF-7_RICTOR<sup>SH</sup> cells on DNMT inhibition by DAC. (<b>H</b>) ChIP-qPCR results (mean ± SEM, <i>n</i> = 3) confirm enhanced binding of ZFX to UGCG promoter in MCF-7_RICTOR<sup>SH</sup> cells on DAC (5 μM) treatment. (<b>I</b>) A schematic representation of pAKT-mediated regulation of histone demethylase KDM5A that regulates UGCG transcription via histone methylation. (<b>J</b>) Immunoblot showing the change in phosphorylation of KDM5A using the pan-phospho-Ser antibody in MCF-7_RICTOR<sup>SH</sup> cells compared to MCF-7 cells. (<b>K</b>, <b>L</b>) Immunoblot (K) and its quantification (mean ± SEM, <i>n</i> = 3) (L) show alterations in KDM5A expression in nuclear and cytoplasmic extracts in MCF-7_RICTOR<sup>SH</sup> cells compared to MCF-7 cells. (<b>M</b>) Immunoblot reveals a decrease in phosphorylation of KDM5A on treatment of MCF-7_ZFX<sup>OE</sup> cells with AKT inhibitor MK2206. (<b>N</b>, <b>O</b>) Results from qRT-PCR (mean ± SEM, <i>n</i> = 3) (<b>N</b>) and immunoblot (O) show increased <i>UGCG</i> expression in MCF-7_RICTOR<sup>SH</sup> cells on KDM5A inhibition. (P) ChIP-qPCR results (mean ± SEM, <i>n</i> = 3) show a reduced H3K4Me3 mark on UGCG promoter in MCF-7_RICTOR<sup>SH</sup> cells that increases on treatment with KDOAM-25 inhibitor (30 μM). Data among two groups were analyzed using an unpaired Student <i><i>t</i></i> test and among multiple groups using One-way ANOVA. <i><i>p</i></i>-value: *<i><i>p</i></i> < 0.05, **<i><i>p</i></i> < 0.01, ***<i><i>p</i></i> < 0.0005, ****<i><i>p</i></i> < 0.0001. Numerical data can be found in <a href="http://www.plosbiology.org/article/info:doi/10.1371/journal.pbio.3003362#pbio.3003362.s015" target="_blank">S4 Data</a>.</p>