RIα<sup>Y122A</sup> disrupts cAMP-stimulated RIα LLPS.

<p><b>(A)</b> Representative maximum intensity projections from confocal Z-stacks showing GFP expression <b>(left)</b> and quantification of puncta per cell <b>(right)</b> from HEK293T cells co-expressing Cα-mCherry plus GFP2-tagged RIα<sup>WT</sup&...

وصف كامل

محفوظ في:
التفاصيل البيبلوغرافية
المؤلف الرئيسي: Ha Neul Lee (7360352) (author)
مؤلفون آخرون: Julia C. Hardy (21781692) (author), Emily H. Pool (14883318) (author), Jin-Fan Zhang (21781695) (author), Su Hyun Kim (828853) (author), William F. Buhl (21781698) (author), Jessica G.H. Bruystens (21781701) (author), Sohum Mehta (2000764) (author), Susan S. Taylor (44167) (author), Jin Zhang (53297) (author)
منشور في: 2025
الموضوعات:
الوسوم: إضافة وسم
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الوصف
الملخص:<p><b>(A)</b> Representative maximum intensity projections from confocal Z-stacks showing GFP expression <b>(left)</b> and quantification of puncta per cell <b>(right)</b> from HEK293T cells co-expressing Cα-mCherry plus GFP2-tagged RIα<sup>WT</sup> (WT, <i>n</i> = 90 cells) or RIα<sup>Y122A</sup> (Y122A, <i>n</i> = 59 cells) before (−) and after (+) Fsk and IBMX stimulation. ****P<i> </i>< 1 × 10<sup>−15</sup>; 2-way ANOVA with Šídák’s multiple comparisons test. <b>(B)</b> Representative fluorescence images and line-intensity profiles of the indicated regions in HEK293T cells expressing smAKAP-EGFP with mRuby2-tagged RIα<sup>WT</sup> or RIα<sup>Y122A</sup>. <b>(C)</b> BRET assay time-course <b>(left)</b> and quantification of Fsk-stimulated change <b>(right)</b> in the GFP2/RLuc8 emission ratio in HEK293T cells co-expressing GFP2-Cα plus RLuc8 fused to either RIα<sup>WT</sup> (<i>n</i> = 3 experiments) or RIα<sup>Y122A</sup> (<i>n</i> = 3 experiments). Untethered RLuc8 co-expressed with GFP2-Cα is used as a control. <i>P</i> = 0.61, unpaired, two-tailed Student <i>t</i> test. Solid lines indicate the mean, and shaded areas indicate SEM. <b>(D)</b> Critical saturation concentration (C-sat) curves of RIα<sup>Y122A</sup>. In vitro phase diagram of RIα<sup>Y122A</sup> vs. PEG 4000 <b>(Left)</b>. Filled circles represent droplets, and empty circles represent no droplets. C-sat curves showing PKA-Cα concentration-dependent LLPS behavior of RIα<sup>WT</sup> (green) and RIα<sup>Y122A</sup> (magenta) <b>(Right)</b>. Dotted lines indicate the boundary between one-phase and two-phase regions (shaded areas). RIα<sup>WT</sup> data were modified from our previous work [<a href="http://www.plosbiology.org/article/info:doi/10.1371/journal.pbio.3003262#pbio.3003262.ref010" target="_blank">10</a>]. <b>(E)</b> Quantification showing basal puncta number per cell in RIα-null MIN6 β cells expressing either mRuby2-tagged RIα<sup>WT</sup> (WT; <i>n</i> = 98 cells) or RIα<sup>Y122A</sup> (Y122A; <i>n</i> = 103 cells). ****<i>P</i> = 5.47 × 10<sup>−12</sup> (WT vs. Y122A); unpaired Komogorov-Smirnov test. Error bars in summary quantification indicate median ± 95% CI. <b>(F)</b> Representative images <b>(left)</b> and quantification of RIα<sup>Y122A</sup> puncta number (<i>n</i> = 53 cells) <b>(middle)</b> and area (for pre-existing puncta) (<i>n</i> = 15 puncta) <b>(right)</b> in RIα-null MIN6 β cells. ns, <i>P</i> = 0.17 for puncta number per cell; ns, <i>P</i> = 0.12 for pre-existing puncta area; paired, two-tailed Student’s <i>t</i> tests. All scale bars, 10 µm. The data underlying this figure can be found in <a href="http://www.plosbiology.org/article/info:doi/10.1371/journal.pbio.3003262#pbio.3003262.s010" target="_blank">S1 Data</a>.</p> <p>(TIF)</p>